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All the zebrafish free swimming movies were analyzed using the image and video processing toolbox of MATLAB. A stepwise image segmentation algorithm was followed. The images were first preprocessed using a customized background subtraction algorithm. Next, the image was thresholded to 1 bit (black/white) to yield a fish outline. A skeletonization algorithm was applied to find the center of the outline, corresponding to the backbone. A cubic spline was then fitted to n points evenly sampling the fish backbone along the backbone arc length. Each step is explained in S1 File and is shown in S1 Fig. 2b1af7f3a8